The Gastropods of Lake Tanganyika,
Diagnostic key, classification, and notes on the fauna

by Kelly West*^1,5, Ellinor Michel*², Jonathan Todd³, David Brown⁴ & Jerod Clabaugh¹

* The first two authors contributed equally. Referencing is legitimate in either order.

¹ Lake Tanganyika Biodiversity Project, Bujumbura, Burundi

² Institute for Biodiversity and Ecosystem Dynamics/Zoological Museum, University of Amsterdam, P.O. Box 94766, 1090 GT Amsterdam, The Netherlands

³ Department of Palaeontology, The Natural History Museum, Cromwell Road, London, SW7 5BD, UK

⁴ Department of Zoology, The Natural History Museum, Cromwell Road, London, SW7 5BD, UK

⁵ IUCN - The World Conservation Union, Eastern Africa Regional Office, P.O. Box 68200, Nairobi 00200, Kenya

Illustrations by Jean-Pierre Kiza & Leonard Papadopoulos

ISBN 9988-0-1821-5

Published in 2003 by the Centre for African Wetlands, Ghana, on behalf of International Association of Theoretical and Applied Limnology, c/o Department of Environmental Sciences, School of Public Health, The University of North Carolina, Chapel Hill, North Carolina 27599-7431 USA, with financial assistance from the World Wide Fund for Nature, Africa and Madagascar Programme which is gratefully acknowledged.

Table of Contents
Preface	7
Acknowledgements	8
Notes for the Reader	9
Introduction to the Fauna	10
Shell Morphology and Identification	11
Diagnostic Keys	22
Taxonomic Classification of Thiarid Gastropods from Lake Tanganyika	25
Thiaridae	25
Genus Anceya 26 Genus Bathanalia 27 Genus Bridouxia 28 Genus Chytra 31 Genus Cleopatra 32 Genus Hirthia 33 Genus Lavigeria, with key to species 35 Genus Limnotrochus 59 Genus Martelia 60 Genus Melanoides 62 Genus Mysorelloides 63 Genus Paramelania 64 Genus Potadomoides 68 Genus Reymondia 69 Genus Spekia 71 Genus Stanleya 73 Genus Stormsia 74 Genus Syrnolopsis 74 Genus Tanganyicia 76 Genus Tiphobia 78 Genus Vinundu 78
Taxonomic Classification of Non-Thiarid Gastropods from Lake Tanganyika	80
Viviparidae	80
Genus Bellamya 80 Genus Neothauma 80
Ampullariidae	81
Genus Pila 81 Genus Lanistes 81
Hydrobiidae	82
Genus Tomichia (?) 83
Bithyniidae	84
Genus Gabbiella 84
Lymnaeidae	84
Genus Lymnaea 84
Ancylidae	84
Genus Burnupia 84 Genus Ferrissia 85
Planorbidae	86
Genus Afrogyrus 86 Genus Ceratophallus 87 Genus Gyraulus 87 Genus Lentorbis 88 Genus Segmentorbis 88 Genus Biomphalaria 90 Genus Bulinus 91
Summary: Updated Checklist of Tanganyikan Thiarid Gastropods	93
Tips for Creating a Collection	97
Glossary	103
Bibliography	117
Index	128
SIL information	131